Morfologia do gênero Baetodes Needham & Murphy (Ephemeroptera: Baetidae) com filogenia do Complexo Baetodes

Abstract

Baetidae is the second richest family of Ephemeroptera, with cosmopolitan distribution and with morphology based on nymphs. The Baetodes Complex groups six Neotropical genera of Baetidae: Baetodes, Lugoiops, Moribaetis, Mayobaetis and Spiritiops which share a posterior seta on tarsal claw. Baetodes, that provides the complex name, is the richest genus among the genera above. Adaptations to climber habit provide to Baetodes an unusual morphology different from no-reophilics genera. Account this and poorly drawn structures, some features of Baetodes create some doubts. On present work, we revised morphology of Baetodes, described new structures, improved generical description, provided a list and images of features with interspecifical variation and, also, proposed the use of new terms. Monophyly of Baetodes Complex was recovered excluding Spiritiops and adding Indobaetis. Prebaetodes and Baetodes were recovered to sister-group, as well Lugoiops and Mayobaetis. The presence of a posterior seta on tarsal claw and the presence of a distomedial projection on article I of forceps, as well as on some former works, were recovered synapomorphic to new Baetodes Complex.